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Encyclopedia / Goldfish & Coldwater / European bitterling
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Variations

1 kinds · tap one for a closer look

European bitterling

Rhodeus amarus · Bitterling

Peaceful and shoaling; males guard mussels in spring.

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8 cmAdult size ✓
UnknownTemperature ?
UnknownpH ?
UnknownSmallest tank ?
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Tank mates
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This one is best kept on its own, or with its own kind.

✕Not recommended
  • European bitterling needs cool water and makes a lot of waste, while Cardinal tetra is tropical.
✕Not recommended
  • European bitterling needs cool water and makes a lot of waste, while Serpae tetra is tropical.
✕Not recommended
  • European bitterling needs cool water and makes a lot of waste, while X-ray tetra is tropical.
Facts checked against sources on 16 Oct 2026. ✓ means a source backs it; ? means no reliable source yet. See sources

Care and water

Hardness ?Unknown
Group size ?6 or more
Temperament ?Peaceful
Swims at ?Middle
Care level ?Moderate
Diet Omnivore
Lifespan ✓1–5 years
Size in shops 2–5 cm
Growth Medium

Behavior

Peaceful and shoaling; males guard mussels in spring.

Check your local laws. Listed (as Rhodeus sericeus amarus) in Annex II of the EU Habitats Directive 92/43/EEC, so EU states must designate conservation areas for it, and in Annex III of the Bern Convention. In Great Britain, where it is introduced, release is regulated under the Wildlife and Countryside Act 1981. The US Fish & Wildlife Service rates its invasion risk to the US as high (a screening, not a ban). Check the rules where you live before you buy one, and never release it into the wild.

  • cool

Food

Small pellets, flake and frozen foods.

Tap a food to see what it is.

Male or female

In spring males take on bright nuptial colours and defend territories around mussels; females grow a long egg-laying tube (ovipositor).

Health

Common problems. Tap one to see the signs. This is a guide, not a diagnosis.

Breeding

Mussel spawner ·

The female uses her ovipositor to lay eggs onto the gills of a live freshwater mussel (Unio or Anodonta) through its exhalant siphon; the male sheds sperm at the inhalant siphon. A tank or pond with live mussels is needed.

Pairing

Group with live mussels.

Setup

Large cool tank with live freshwater mussels and plants.

What starts it

Spring warm-up.

Eggs

One to six eggs per laying, about 80-300 per female per season. Eggs hatch in 24-36 hours, and the young stay in the mussel for 3-4 weeks before swimming out at about 10 mm.

Raising the fry

Fry leave the mussel after about 4 weeks and feed on infusoria.

Common mistakes
  • No mussels
  • Warm water
Full breeding guide, day-by-day plan and simulator →

Where it lives in the wild

Loading the map…
Native range (rough area)Centre of the range
RegionEurope – France east to the Black Sea basin
Rivers and lakesRivers of central and eastern Europe, including the Danube basin
HabitatSlow, weedy rivers, ponds and lakes with mussels
Native rangeAbout 1,500 km across
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The story of the european bitterling

A small European fish famous for laying its eggs inside living freshwater mussels; it cannot breed without a mussel host. It was once used in human pregnancy tests.

Marcus Bloch described this fish in 1782 as Cyprinus amarus. It is famous for its breeding behaviour: females lay eggs inside freshwater mussels using a long ovipositor. The young develop inside the mussel before swimming out.

First described by Marcus Elieser Bloch in 1782. Named for 'amarus' means 'bitter'.

  • It lays its eggs inside living freshwater mussels.
  • Females grow a long egg-laying tube during the breeding season.
  • Males turn pinkish-purple when breeding.

Sources

12 facts checked with sources

  • Scientific name

    Older literature (and the EU Habitats Directive annex) calls it Rhodeus sericeus amarus; R. sericeus is now the Amur bitterling of East Asia. Shops may also sell the East Asian rosy bitterling R. ocellatus as "bitterling". Coldwater fish.

    Eschmeyer's Catalog of Fishes: Rhodeus amarus (online version updated 21 Sep. 2026) · FishBase: Rhodeus amarus summary (main ref. Kottelat & Freyhof 2007)
  • Familycorrected

    Legacy has no family; value is new. Formerly a subfamily of Cyprinidae.

    FishBase: Rhodeus amarus summary (main ref. Kottelat & Freyhof 2007)
  • Size

    size-pick: TL. Common length 5 cm, maximum record 11.2 cm. Legacy 8 cm is a large adult; no Seriously Fish profile.

    FishBase: Rhodeus amarus summary (main ref. Kottelat & Freyhof 2007) · Wikipedia: European bitterling (cites FishBase; Tarkan et al. 2005)
  • Temperature

    No keeping range found. A temperate fish that spawns from about 10-15 °C up, optimum 23 °C; legacy top of 22 °C is plausible.

    US Fish & Wildlife Service: Ecological Risk Screening Summary, European Bitterling (Rhodeus amarus) · Croatian Agency for Environment and Nature (HAOP): Monitoring programme for Rhodeus amarus (government PDF)
  • Lifespancorrected

    Most wild fish live 1-2 years (mature at 1 year); 5 years is exceptional. Captive lifespan not sourced. Legacy 4-5 years is the top end.

    FishBase: Rhodeus amarus summary (main ref. Kottelat & Freyhof 2007)
  • Swimming level

    Benthopelagic; no source says mid-water in substance.

    FishBase: Rhodeus amarus summary (main ref. Kottelat & Freyhof 2007)
  • Where it livescorrected

    Legacy has no range; value is new. Introduced to Britain and Italy.

    FishBase: Rhodeus amarus summary (main ref. Kottelat & Freyhof 2007)
  • Male or female

    Legacy also mentions male tubercles; not checked.

    US Fish & Wildlife Service: Ecological Risk Screening Summary, European Bitterling (Rhodeus amarus)
  • Breeding

    Spawns late April to August.

    US Fish & Wildlife Service: Ecological Risk Screening Summary, European Bitterling (Rhodeus amarus) · Croatian Agency for Environment and Nature (HAOP): Monitoring programme for Rhodeus amarus (government PDF)
  • Eggs and youngcorrected

    Legacy "few eggs per female" understates the seasonal total; legacy 4 weeks in the mussel is right.

    Croatian Agency for Environment and Nature (HAOP): Monitoring programme for Rhodeus amarus (government PDF) · Wikipedia: European bitterling (cites FishBase; Tarkan et al. 2005)
  • StoryFishBase: Rhodeus amarus summary (main ref. Kottelat & Freyhof 2007)
  • traits.legalcorrected

    Annex II means habitat protection, not a keeping ban; check national rules before collecting wild fish. The host mussels (Unio, Anodonta) are protected in some countries, but no source was checked for that.

    Croatian Agency for Environment and Nature (HAOP): Monitoring programme for Rhodeus amarus (government PDF) · US Fish & Wildlife Service: Ecological Risk Screening Summary, European Bitterling (Rhodeus amarus)